Gobies are fish of the family Gobiidae, one of the largest fish families comprising more than 2,000 species in more than 200 genera. Most of them are relatively small, typically less than in length. Gobies include some of the smallest vertebrates in the world, such as Trimmatom nanus and Pandaka pygmaea, which are under long when fully grown. Some large gobies, such as some species of the genera Gobioides or Periophthalmodon, can reach over in length, but that is exceptional. Generally, they are benthic, or bottom-dwellers. Although few are important as food for humans, they are of great significance as prey species for commercially important fish such as cod, haddock, sea bass, and flatfish. Several gobies are also of interest as aquarium fish, such as the bumblebee gobies of the genus Brachygobius. Phylogenetic relationships of gobies have been studied using molecular data.